بیماری‌هامراقبت‌های بهداشتیپزشکی و سلامت

پاسخ ایمنی هومورال (تولید آنتی بادی) بواسطه لنفوسیت B Humoral immune response (antibody production) by B lymphocytes

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ایمنیم هوققووال ‎ear aE)‏ اس ۲ 8 ‎a‏ اعم ‎Hummora‏ 3+3 : 3 ل بیس ‎«(an‏ »> ۵ ال ' عل 5 ۴ یک ۳ ۰ ِ اس > 07 — eh SS) 9 (۹ ‎ay — e‏ عه ‏35 - يبيد ‏ا ‏۰ ‏.

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00 نفوسیت 8 چگونه با تشوسیت ‎oe‏ اه سيت پا ۱ را تولید می کنند؟

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ee Ter ae Re hover oi ore Soccer eee leer 9 حدف از بوجود آمدن پاسخ ایمنی هومورال ... . . . . . . می‌باشد.

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Immature B cells

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‎a‏ و ‎۵ 6 6 ‎2 ‎Di Yip) ‏ااا‎ ‎Surrogate Surrogate ‏ها‎ > Vpre Bands Vpre'8andAs ‎Surogate ‎Aight chain ‏دنت ‏— ‎STEMCELL ‎Meehan genes Germtine ‎Germtine‏ ممصمو مامز ‎00 ‎rar ‎Membrane 1 Heavy chain Tight hain ‏يلاس ‏ب ا 19509 تس ‎cas ‘cos, ‏مسب لواو‎ ‎

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Follicul B2B cell “ا وی immature B

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Conventional | Marginal zone eee ace B-2 cells Bells ‘When first produced Fetus After birth After birth \N-tegions in VDJ junctions. Few Extensive Yes Primary lean ee ee 55 ‏پم ای مور‎ [| ۳ || Longives ‏سا سا س | رس‎ Isotypes secreted IgM >> IgG IgG > IgM IgM > IgG ‏ال ال تسه‎ Response to protein antigen Maybe Yes Yes Requirement for T-cell help No Yes Sometimes ‘Somatic hypermutation ‘Low-none High 2 ‘Memory development Little or none Yes 7 FE OE REE Sey

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ao ‏تسسا اس تا‎ response B2 cells mediated immune response to TD-Ag* Ba cells mediated immune response to TI-Ag*

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B cell antigen recognition The B cell antigen receptor is cell surface immunoglobulin (smlg) AN 4 N Soluble antibodies and surface membrane immunoglobulin bind directly to native antigens

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Oe ume gat cd و1 مذو سبابيجمن Caer eat eo Pee

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B cells recognize antigen BCRs directly recognize the epitope on the surface of Ag* No APC + ۳0 ۱/۳۵۵۵ ادك ل مره مورک

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Extracellular

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B-cell coreceptor C3d SSS — CR2 (CD21) cD19 TAPA-1 (CD81) Others?

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Ig-a/Ig-B B-cell membrane Cytoplasm ITAM sequence erate Sa

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(a) Tl-1 antigen (b) TD antigen Ty cell

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۱ ‏ا‎ ‎seed Ah es Ach ite Memory T cell

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3 a 3 4 15 9 aa ۱ 3 if He) memory cells

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9 5 nin serum, Primary Secondary response response units per ml & 2 5 g z 9 8 > 3 3 2 3 < 2° Ag Time after immunization

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Tule But ‏ارزو‎ | longer | ‏میسن ن‎ ا يتك

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لنفوسيت هاى 8 جكونه انواع كلاس آنتى بادى را توليد مى نمايند؟ شكل كيرى ياسخ ايمنى هومورال در مركز زايا (زرمينال سئتز)؟ 0ل ا انا ل ال روز eC Re CNL eI ro BOR ae kerr cc 0

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Afferent lymphatic Paracortex Initial T-cell and B-cell | [activation Cortex Medulla Plasma-cell secretion of antibody 0 Primary follicle B-cell activation Germinal center B-cell proliferation and differentiation Secondary follicle Efferent lymphatic vessel

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Follicle Light zone Germinal center

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ار یت ۸:19 دا ك2 اكه )6ه دو 59 Grote RECOMBINATION ۳ لس يردن

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Differentiation of B cells in germinal Center ۱ ‏م‎ ee ‏سلولهای 3] در تولید آنتی بادی بوجود می آید.‎ affinity maturation

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9 centrocytes byt ‘ining fo Ag-Ab compl

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Recognition Activation phase: B cell ee proliferation and differentiation 0 © 0 Hel J tis, Antibody ‏)مانا‎ T cells (Bere Clonal secretion|| cancion] ‏سس‎ ‎ee ‘expressing ‏وا‎ ‎B cell 3 Isotype ۰ (0 < ©1197“ | Antigen Resting Activated rit va Affinity || mature B cel 2 2 Bs maturation High-affinity * ‏موب‎ ‎‘expressing B 4 A affinity IgG Memory B cell | Figure 9-1 Phases of the humoral immune response. The activation of 8 cells is initiated by specific recognition of antigens by the surface Ig receptors of the cells, Antigen and other stimuli, inciuding helper T cells stimulate the proliferation and ditter- Entiation of the specific B cell clone. Progeny of the clone may produce lgM or other Ig isotypes (e.9., i, oF may persist as memory cells

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nl ۱۳۸ - 06 9 'oG3 ge gf cytokines B-cell activation TGF-B = & lgG2b هوا مار > او © کرت

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Helper Teel pao. eso. ‏و‎ ۱ ‏تست‎ 2 ‘Aotivated 2 Mucosal tissues, cytokines TEN-y] [IL-4 (e.g., TGF-B) 19G subclasses IgM ie Igeay ‘ge 9۸ ‘Complement Fe receptor- Immunity Mucosal | setivation dependent against immunity phagocyte Rolminths Gransport of Fosponaes: IgA through complement Mast cell Spinel} activation: Gearanulat Reonatal Immunity | {immeckat Transtor) | hyporseneitivity) Figure 9-15 1g heavy chain tkotype switching. 8 cells activated by helper T cell signals (CD40L, cytokines) undergo switching to different Ig iso ‘types, which mediate distinct effector functions. Selected examples of switehed isotypes in hurmans are shawn

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Naive B cell “antigen VDI Sy Gi CR by Sy Cy le Se Ce ERROR ‏ننه © اه هات‎ HOLE 01 ‏ته‎ / -3 No signals fram| Signals fom helper T-cells helper T cells (E40 ligand. cytokines) Garmiine 7 1 Garmiina © esis ccs eee | Geta ee nec | S| ‏الممل هه ابسو‎ 7 ۷ ‏کی‎ See vos VW cy Sece voc, om ‏مهم‎ y MRNA + ‎oy protein‏ سس ‎ ‎ ‎ ‎| activated 6 ce ‎ ‎ig example, ge). These sur ‎ ‎Transcription ‎through yore Sud ‎Recombination ‎of Sy with ‎oF Se dete ‎3t interven ‎© genes 4 ‎ ‎Transcnpuon: ‎FINA splicing ‎ ‎ ‎ ‎“he absence of helper Teel signals, Bcelke prackice ight ‏که‎ Reiner chat agnis ‏در و امد‎ ais 2 cn Ui ‎Translation ‎] Site for serine eranccriptonh Chats that ere she swipe Cy yerien located between re Lees me ey ‎ ‎ ‎ ‎ ‎ ‎ ‎ ‎ ‎ ‎

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Plasma cell Proliferating B cells (centrocytes) Differentiation cytokines: IL-2, IL-4, IL-5, IFN~y< TGF-B| Activated B cell (centroblast) Proliferation cytokines: IL-2, 1L-4, IL-5 ی میت رک

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